Silvicultural and mensurational structure of Pinus nigra Stands in Ukraine based on the 2011 state forest inventory data

Yurii Siruk*, Iryna Siruk, Oleg Prut

qarpofor@gmail.com

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Abstract

The study aimed to identify the territorial, forest-site, silvicultural, and mensurational patterns governing the formation of Crimean pine and Austrian pine stands in Ukraine. The analysis was based on a compartment-level database of the State Forest Inventory as of 2011. Structured queries in the Microsoft SQL Server database management system, weighted mean calculations, and regression analysis of age-related changes in stand mensurational characteristics were applied. The total area of the studied stands amounted to 134.1 thousand ha, of which 99.8% was occupied by Crimean pine (P. nigra var. pallasiana) and 0.2% by Austrian pine (P. nigra var. austriaca). Most Crimean pine stands were concentrated in the southern and eastern regions of Ukraine, whereas Austrian pine stands had a local distribution, predominantly in the western part of the country. Very dry and dry forest-site conditions accounted for 84.8% of the total area of Crimean pine stands, primarily in dry fairly fertile site types and dry poor site types. Austrian pine occurred more frequently in fresh fairly fertile and fertile site types. Artificially established and pure stands predominated in the structure of both taxa. Under dry site conditions, artificially established stands accounted for 91.5% of Crimean pine stands and 98.5% of Austrian pine stands. The highest site-quality classes, mean heights, mean diameters, growing stock volumes, and proportions of merchantable timber were generally recorded in fairly fertile and fertile site types. By contrast, the productive potential of stands growing in dry poor site types was constrained by moisture deficiency and low soil fertility. The diversity of associated tree species increased with increasing site trophicity: P. sylvestris predominated in poor site types, whereas the contribution of broadleaved hardwood species increased in fairly fertile and fertile site types

Keywords

Crimean pine; Austrian pine; forest-site types; growing stock volume; productivity; merchantable timber; growth dynamics

  1. Adamenko, S., Shlapak, V., Kozachenko, I., & Parubok, M. (2018). Peculiarities of seasonal growth of Pinus nigra J.F. Arnold under the conditions of introduction in the Right-Bank Forest-Steppe of Ukraine. Journal of Forest Science, 64(8), 340-344. doi: 10.17221/63/2018-JFS.
  2. Ayan, S., Yücedağ, C., & Simovski, B. (2021). A major tool for afforestation of semi-arid and anthropogenic steppe areas in Turkey: Pinus nigra J.F. Arnold subsp. pallasiana (Lamb.) Holmboe. Journal of Forest Science, 67(10), 449-463. doi: 10.17221/74/2021-JFS.
  3. Bessonova, V., & Yakovlieva-Nosar, S. (2022). Dynamics of non-structural carbohydrates in Pinus pallasiana D. Don needles under different forest growth conditions of ravine anti-erosion plantations. Folia Forestalia Polonica, 64(1), 38-48. doi: 10.2478/ffp-2022-0004.
  4. Bessonova, V., & Yusypiva, T. (2021). Physiological parameters of the state of Pinus pallasiana D. Don in different forest-growth conditions in Ravine Viyskovyi. Ekológia (Bratislava), 40(2), 137-144. doi: 10.2478/eko-2021-0016.
  5. Bricca, A., Zerbe, S., Fellin, H., Alessi, N., Maccherini, S., & Bonari, G. (2025). Passive rewilding of old-established plantations into native forests. Journal of Vegetation Science, 36(6), article number e70085. doi: 10.1111/jvs.70085.
  6. Bulut, S., Günlü, A., & Şatır, O. (2023). Estimating net primary productivity of semi-arid Crimean pine stands using biogeochemical modelling, remote sensing, and machine learning. Ecological Informatics, 76, article number 102137. doi: 10.1016/j.ecoinf.2023.102137.
  7. Candel-Pérez, D., Lucas-Borja, M.E., García-Cervigón, A.I., Tíscar, P.A., Andivia, E., Bose, A.K., Sánchez-Salguero, R., Camarero, J.J., & Linares, J.C. (2022). Forest structure drives the expected growth of Pinus nigra along its latitudinal gradient under warming climate. Forest Ecology and Management, 505, article number 119818. doi: 10.1016/j.foreco.2021.119818.
  8. Collado, E., De Cáceres, M., de-Dios-García, J., Rubio-Gutiérrez, R., Roženbergar, D., & Piqué, M. (2026). Forest management and drought interaction on tree growth in black pine forests across the Mediterranean Basin. Dendrochronologia, 96, article number 126497. doi: 10.1016/j.dendro.2026.126497.
  9. Devi, N.M., Kukarskih, V.V., & Bubnov, M.O. (2023). Elevational changes in climate response of Pinus nigra pallasiana tree-rings on the Crimean Peninsula. Dendrochronologia, 79, article number 126092. doi: 10.1016/j.dendro.2023.126092.
  10. Dias, A., Carvalho, A., Silva, M.E., Lima-Brito, J., Gaspar, M.J., Alves, A., Rodrigues, J. C., Pereira, F., Morais, J., & Lousada, J.L. (2020). Physical, chemical and mechanical wood properties of Pinus nigra growing in Portugal. Annals of Forest Science, 77, article number 72. doi: 10.1007/s13595-020-00984-8.
  11. Dias, A., Louzada, J., & Fonseca, T. (2022). Growth pattern of European black pine outside its current natural range: A case study in Portugal. Land, 11(6), article number 794. doi: 10.3390/land11060794.
  12. Giovannelli, G., Tallieu, C., Davi, H., Guibal, F., Scotti-Saintagne, C., & Fady, B. (2026). Adaptive effects of geographic origin on ring-width and radial growth-climate relationships in Pinus nigra Arnold, the European black pine. Annals of Forest Science, 83, article number 31. doi: 10.1186/s13595-026-01347-5.
  13. Gülsoy, S., & Çinar, T. (2019). The relationships between environmental factors and site index of Anatolian black pine (Pinus nigra Arn. subsp. pallasiana (Lamb.) Holmboe) stands in Demirci (Manisa) district, Turkey. Applied Ecology and Environmental Research, 17(1), 1235-1246. doi: 10.15666/aeer/1701_12351246.
  14. Information from the State Forest Cadastre. (2019). State forest cadastre as of 1 January 2011: General data on the forest fund of Ukraine by administrative-territorial units. Retrieved from https://data.gov.ua/dataset/341e5bd6-3855-4507-9a53-f95a9a1e3035.
  15. Ivetić, V., Tsakaldimi, M., Ganatsas, P., Kerkez Janković, I., & Devetaković, J. (2021). Freezing and heating tolerance of Pinus nigra seedlings from three south to north Balkan provenances. Sustainability, 13(16), article number 9290. doi: 10.3390/su13169290.
  16. Izmir, Ş.C., Jevšenak, J., Krajnc, L., Hafner, P., & Köse, N. (2024). Distinct spatial patterns in climate-growth relationships, vegetation and resilience indices of black pine (Pinus nigra J. F. Arnold) from its northern and southern distribution range. Dendrochronologia, 88, article number 126236. doi: 10.1016/j.dendro.2024.126236.
  17. Koval, I. (2013). Climatic signal in earlywood, latewood and total ring width of Crimean pine (Pinus nigra subsp. pallasiana) from Crimean Mountains, Ukraine. Baltic Forestry, 19(2), 245-251.
  18. Krasnov, V.P., Orlov, O.O., & Vedmid, M.M. (2009). Atlas of indicator plants and forest site types of Ukrainian Polissia. Zvyagel: NOVOhrad.
  19. Lucas-Borja, M.E., Jing, X., Candel-Perez, D., Parhizkar, M., Rocha, F., Heydari, M., Muñoz-Rojas, M., & Zema, D.A. (2022). Afforestation with Pinus nigra Arn ssp. salzmannii along an elevation gradient: Controlling factors and implications for climate change adaptation. Trees, 36, 93-102. doi: 10.1007/s00468-021-02184-x.
  20. Martín-Alcón, S., & Coll, L. (2016). Unraveling the relative importance of factors driving post-fire regeneration trajectories in non-serotinous Pinus nigra forests. Forest Ecology and Management, 361, 13-22. doi: 10.1016/j.foreco.2015.11.006.
  21. Nagel, T.A., & Cerioni, M. (2023). Structure and dynamics of old-growth Pinus nigra stands in Southeast Europe. European Journal of Forest Research, 142, 537-545. doi: 10.1007/s10342-023-01540-5.
  22. Olsson, S., Grivet, D., Cattonaro, F., Vendramin, V., Giovannelli, G., Scotti-Saintagne, C., Vendramin, G.G., & Fady, B. (2020). Evolutionary relevance of lineages in the European black pine (Pinus nigra) in the transcriptomic era. Tree Genetics & Genomes, 16, article number 30. doi: 10.1007/s11295-020-1424-8.
  23. Pluhatar, Y.V., Shvets, Y.P., Trofymenko, I.O., & Diahovets, O.O. (2010). Crimean pine: Range, forest types, and productivity. Scientific Bulletin of UNFU, 20(6), 39-50.
  24. Raptis, D.I., Kazana, V., Kazaklis, A., & Stamatiou, C. (2021). Mixed-effects height–diameter models for black pine (Pinus nigra Arn.) forest management. Trees, 35, 1167-1183. doi: 10.1007/s00468-021-02106-x.
  25. Seki, M., & Sakici, O.E. (2022). Ecoregional variation of Crimean pine (Pinus nigra subspecies pallasiana [Lamb.] Holmboe) stand growth. Forest Science, 68(5-6), 452-463. doi: 10.1093/forsci/fxac030.
  26. Skrobach, T.B. (2006). Black pine (Pinus nigra Arn.) in forest stands of the Western region of Ukraine. (PhD dissertation, Ukrainian National Forestry University, Lviv, Ukraine).
  27. Soroka, M., & Yuskevych, T. (2011). Phytosociological aspects of the naturalisation of black pine (Pinus nigra Arn.). Proceedings of the Forestry Academy of Sciences of Ukraine, 9, 58-63.
  28. Tereshchenko, L.I., Los, S.A., Petrenko, M.I., & Samoday, V.P. (2019). Productivity and condition of Austrian pine (Pinus nigra var. austriaca Asch. et Gr.) in central and north-eastern parts of Ukraine. Forestry and Forest Melioration, 135, 68-78. doi: 10.33220/1026-3365.135.2019.68.
  29. Tudoran, G.-M., Cicșa, A., Dobre, A.-C., Cicșa, M., Pascu, I.-S., & Leca, Ş. (2023). Health and growth of black pine outside its natural distribution range in the Romanian Carpathians. Forests, 14(5), article number 884. doi: 10.3390/f14050884.
  30. Ukrderzhlisproekt Production Association. (2006). Instruction for forest management planning of the forest fund of Ukraine. Part 1. Field works. Irpin: Ukrderzhlisproekt.
  31. Ukrderzhlisproekt Production Association. (2012). Directory on the forest fund of Ukraine based on the state forest inventory as of January 1, 2011. Irpin: Ukrderzhlisproekt.
  32. Vacek, Z., Cukor, J., Vacek, S., Gallo, J., Bažant, V., & Zeidler, A. (2023). Role of black pine (Pinus nigra J.F. Arnold) in European forests modified by climate change. European Journal of Forest Research, 142, 1239-1258. doi: 10.1007/s10342-023-01605-5.
  33. Versace, S., et al. (2022). New evidence for population-specific responses to drought events from tree ring chronologies of Pinus nigra ssp. laricio across the entire distribution range. Agricultural and Forest Meteorology, 323, article number 109076. doi: 10.1016/j.agrformet.2022.109076.
  34. Yuskevych, T.V., Skrobach, T.B., & Smikh, V.M. (2011). Distribution of Pinus nigra Arn. and Pinus pallasiana D. Don in forest stands of the Western region of Ukraine. Scientific Bulletin of UNFU, 21(17), 74-76.
  35. Zeidler, A., Borůvka, V., Tomczak, K., Vacek, Z., Cukor, J., Vacek, S., & Tomczak, A. (2024). The potential of non-native pines for timber production – a case study from afforested post-mining sites. Forests, 15(8), article number 1388. doi: 10.3390/f15081388.
Siruk, Yu., Siruk, I., & Prut , O. (2026). Silvicultural and mensurational structure of Pinus nigra Stands in Ukraine based on the 2011 state forest inventory data. Scientific Horizons, 29(3), 28-43. https://doi.org/10.48077/scihor3.2026.28